Seed dispersal by animals is often used as a classical example of animal-plant mutualism (Snow 1971, Mckey 1975, van der Pijl 1983). Although this interaction has received much attention (Wheelwright and Orians 1982, Herrera 1985, Howe 1989), the consequences of seed dispersal on plant demography and evolution remain poorly understood (Howe 1989, Jordano 1992).
